The origin and subsequent evolution of land plants transformed our planet utterly. These changes were brought about by key innovations in the land plant lineage, the correct sequence of which is critical to understanding the physiological and developmental evolution that underpin plant evolutionary history. Early land plant fossil remains occur as dispersed spores and compressions of whole plants that inform gross anatomy. The best known record of detailed anatomy is provided by the late Devonian Rhynie Chert, but an older and more diverse record occurs in charcoal lagerstätte from the lower Devonian of Brown Clee Hill, Shropshire. Charcolification produces chemically ...
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